Humans are remarkably similar to other apes. Like us, chimpanzees and orangutans are extremely clever, use tools and exhibit rudimentary understanding of causality and what others intend. However, other apes are not nearly as good at understanding the intentions of others nor nearly so eager to accommodate or help them. By contrast, right from an early age, humans are eager to help and share. It was this combination of understanding what others intend along with impulses to help and please them that enabled our ancestors to coordinate behavior in pursuit of common goals—with spectacular consequences later on. So how and why did such other-regarding capacities emerge in creatures as self-serving as non-human apes are? And why did they emerge...